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Al-Quso, 37, was on the FBI’s most wanted list, with a $5 million reward for information leading to his capture. He was indicted in the U.S. for his role in the 2000 bombing of the USS Cole in the harbor of Aden, Yemen, in which 17 American sailors were killed and 39 injured.

Al-Quso was believed to have replaced Anwar al-Awlaki as the group’s head of external operations. Al-Awlaki was killed in a U.S. airstrike last year.

The new Yemeni president, Abed Rabbo Mansour Hadi, has promised improved cooperation with the U.S. to combat the militants. On Saturday, he said the fight against al Qaeda was in its early stages. Hadi took over in February from longtime authoritarian leader Ali Abdullah Saleh.