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Both mistakes happened in the span of six plays in Minnesota’s 24-13 upset of the 49ers.

“My interpretation of it was that he could do that based upon the time factors and not knowing it was a challengeable play to begin with when he called timeout,” Roan said. “If you don’t have a timeout to lose, you can’t make a challenge.”

Earlier Sunday, the NFL players’ union sent an open letter to team owners calling for an end to the lockout.

In the Lions-Titans and Bengals-Redskins games, officials marched off too much yardage on penalties.

Lions linebacker Stephen Tulloch’s helmet-to-helmet hit on Craig Stevens wound up as a 27-yard penalty in Tennessee’s 44-41 overtime win. In OT, from the Titans 44, Jake Locker passed to Stevens over the middle for a 24-yard gain and Tulloch was flagged for the hit. Fourteen yards were added to the end of the play, which then was reviewed and overturned because the ball hit the ground.

However, the penalty still is enforced. Instead of 15 yards, officials marked it off from the Detroit 44 _ the wrong spot.

“As soon as the play was declared incomplete it becomes a first down and it becomes 15 yards from the play before,” Lions coach Jim Schwartz said.

The Redskins were penalized 20 yards instead of 15 for unsportsmanlike conduct in the final seconds of their 38-31 loss.

Robert Griffin III spiked the ball to stop the clock with 7 seconds left. Then tight end Fred Davis was called for a 5-yard false start penalty.

According to Washington coach Mike Shanahan, at least one official indicated there would be a 10-second runoff, ending the game _ and the Bengals, led by coach Marvin Lewis, started walking onto the field. There shouldn’t have been a runoff, though, because the clock had been stopped by the spike. The Redskins began arguing, and eventually the unsportsmanlike conduct penalty was called.

The officials never announced specifically who the call was against, just that the penalty would be added to the false start, a total of 20 yards. But they walked off 25 yards _ the official game play-by-play said 20 yards were enforced for the unsportsmanlike conduct.

That left the Redskins with a third-and-50.

“They threw the flag at us, and there was half of the (Bengals) team on the field,” Shanahan said. “I was disappointed in that.”

The players’ union posted an open letter to team owners calling on them to end the lockout of the regular officials that began four months ago. The NFL used replacements in 2001 for one week before a new deal was reached.

This year, criticism from coaches and players has mounted for the replacements, who come from lower college levels or from other leagues such as Arena Football.

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