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Williams’ older sister Venus won Monday, her first victory at Wimbledon since 2011. But there were losses for 2011 U.S. Open champion Samantha Stosur, 18th-seeded Sloane Stephens and 27th-seeded Garbine Muguruza, the 20-year-old Spaniard who surprisingly beat Serena Williams in the second round of the French Open last month.

Three lower-seeded men lost, too: No. 18 Fernando Verdasco, No. 25 Andrea Seppi and No. 31 Vasek Pospisil.

Stephens had reached at least the fourth round at six consecutive Grand Slam tournaments, the longest active streak for a woman, but she ran up against a former top-10 player in Maria Kirilenko and was beaten 6-2, 7-6 (6).

Muguruza, meanwhile, saved a hard-to-believe 12 match points but couldn’t erase No. 13, double-faulting twice in a row to get edged 6-3, 3-6, 7-5 by 22-year-old CoCo Vandeweghe, an American ranked 51st.

“It was crazy at the end,” Vandeweghe said.